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Biology

Biology

Biology is the study, or science, of life. It is concerned with the characteristics and behaviors of organisms, how species and individuals come into existence, and the interactions they have with each other and with the environment. Biology encompasses a broad spectrum of academic fields that are often viewed as independent disciplines. However, together they address the phenomenon of life over a wide range of scales. At the atomic and molecular scale, life is studied in the disciplines of molecular biology, biochemistry, and molecular genetics. At the level of the cell, it is studied in cell biology, and at multicellular scales, it is examined in physiology, anatomy, and histology. Developmental biology studies life at the level of an individual organism's development or ontogeny. Moving up the scale towards more than one organism, genetics considers how heredity works between parent and offspring. Ethology considers group behavior of more than one individual. Population genetics looks at the level of an entire population, and systematics considers the multi-species scale of lineages. Interdependent populations and their habitats are examined in ecology and evolutionary biology. A speculative new field is astrobiology (or xenobiology), which examines the possibility of life beyond the Earth.
Biology studies the variety of life (clockwise from top-left) E. coli, tree fern, gazelle, Goliath beetle

Principles of biology

Unlike physics, biology does not usually describe systems in terms of objects which obey immutable physical laws described by mathematics. Nevertheless, the biological sciences are characterized and unified by several major underlying principles and concepts: universality, evolution, diversity, continuity, homeostasis, and interactions.

Universality: Biochemistry, cells, and the genetic code

mathematics]] Main articles: Life The most salient example of biological universality is that all living things share a common carbon-based biochemistry and in particular pass on their characteristics via genetic material, which is based on nucleic acids such as DNA and which uses a common genetic code with only minor variations. Another universal principle is that all organisms (that is, all forms of life on Earth except for viruses) are made of cells. Similarly, all organisms share common developmental processes. For example, in most metazoan organisms, the basic stages of early embryonic development share similar morphological characteristics and include similar genes.

Evolution: The central principle of biology

Main article: Evolution The central organizing concept in biology is that all life has a common origin and has changed and developed through the process of evolution (see Common descent). This has led to the striking similarity of units and processes discussed in the previous section. Charles Darwin established evolution as a viable theory by articulating its driving force, natural selection (Alfred Russell Wallace is recognized as the co-discoverer of this concept). Genetic drift was embraced as an additional mechanism of evolutionary development in the modern synthesis of the theory. The evolutionary history of a species— which describes the characteristics of the various species from which it descended— together with its genealogical relationship to every other species is called its phylogeny. Widely varied approaches to biology generate information about phylogeny. These include the comparisons of DNA sequences conducted within molecular biology or genomics, and comparisons of fossils or other records of ancient organisms in paleontology. Biologists organize and analyze evolutionary relationships through various methods, including phylogenetics, phenetics, and cladistics (The major events in the evolution of life, as biologists currently understand them, are summarized on this evolutionary timeline).

Diversity: The variety of living organisms

evolutionary timeline, based on rRNA gene data, showing the separation of the three domains bacteria, archaea, and eukaryotes as described initially by Carl Woese. Trees constructed with other genes are generally similar, although they may place some early-branching groups very differently, presumably owing to rapid rRNA evolution. The exact relationships of the three domains are still being debated.]] Despite its underlying unity, life exhibits an astonishingly wide diversity in morphology, behavior, and life histories. In order to grapple with this diversity, biologists attempt to classify all living things. Scientific classification seeks to reflect the evolutionary trees (phylogenetic trees) of the organism being classified. Classification is the province of the disciplines of systematics and taxonomy. Taxonomy places organisms in groups called taxa, while systematics seeks to define their relationships with each other. This clasification technique has evolved to reflect advances in cladistics and genetics, shifting the focus from physical similarities and shared characteristics to phylogenetics. Traditionally, living things have been divided into five kingdoms: :Monera -- Protista -- Fungi -- Plantae -- Animalia However, many scientists now consider this five-kingdom system to be outdated. Modern alternative classification systems generally begin with the three-domain system: :Archaea (originally Archaebacteria) -- Bacteria (originally Eubacteria) -- Eukaryota These domains reflect whether the cells have nuclei or not, as well as differences in the cell exteriors. There is also a series of intracellular parasites that are progressively "less alive" in terms of metabolic activity: :Viruses -- Viroids -- Prions

Continuity: The common descent of life

Main article: Common descent Up into the 19th century, it was commonly believed that life forms could appear spontaneously under certain conditions (see abiogenesis). This misconception was challenged by William Harvey's diction that "all life [is] from [an] egg" (from the Latin "Omne vivum ex ovo"), a foundational concept of modern biology. It simply means that there is an unbroken continuity of life from its initial origin to the present time. A group of organisms is said to share a common descent if they share a common ancestor. All organisms on the Earth have been and are descended from a common ancestor or an ancestral gene pool. This last universal common ancestor of all organisms is believed to have appeared about 3.5 billion years ago. Biologists generally regard the universality of the genetic code as definitive evidence in favor of the theory of universal common descent (UCD) for all bacteria, archaea, and eukaryotes (see: origin of life).

Homeostasis: Adapting to change

Main article: Homeostasis Homeostasis is the ability of an open system to regulate its internal environment to maintain a stable condition by means of multiple dynamic equilibrium adjustments controlled by interrelated regulation mechanisms. All living organisms, whether unicellular or multicellular, exhibit homeostasis. Homeostasis manifests itself at the cellular level through the maintenance of a stable internal acidity (pH); at the organismic level, warm-blooded animals maintain a constant internal body temperature; and at the level of the ecosystem, as when atmospheric carbon dioxide levels rise and plants are theoretically able to grow healthier and remove more of the gas from the atmosphere. Tissues and organs can also maintain homeostasis.

Interactions: Groups and environments

organ of the genus Amphiprion that dwell among the tentacles of tropical sea anemones. The territorial fish protects the anemone from anemone-eating fish, and in turn the stinging tentacles of the anemone protects the anemone fish from its predators]] Every living thing interacts with other organisms and its environment. One reason that biological systems can be difficult to study is that so many different interactions with other organisms and the environment are possible, even on the smallest of scales. A microscopic bacterium responding to a local sugar gradient is responding to its environment as much as a lion is responding to its environment when it searches for food in the African savannah. For any given species, behaviors can be co-operative, aggressive, parasitic or symbiotic. Matters become more complex when two or more different species interact in an ecosystem. Studies of this type are the province of ecology.

Scope of biology

Main article: List of biology disciplines Biology has become such a vast research enterprise that it is not generally regarded as a single discipline, but as a number of clustered sub-disciplines. This article considers four broad groupings. The first group consists of those disciplines that study the basic structures of living systems: cells, genes etc.; the second group considers the operation of these structures at the level of tissues, organs, and bodies; the third group considers organisms and their histories; the final constellation of disciplines focuses on their interactions. It is important to note, however, that these boundaries, groupings, and descriptions are a simplified characterization of biological research. In reality, the boundaries between disciplines are fluid, and most disciplines frequently borrow techniques from each other. For example, evolutionary biology leans heavily on techniques from molecular biology to determine DNA sequences, which assist in understanding the genetic variation of a population; and physiology borrows extensively from cell biology in describing the function of organ systems.

Structure of life

DNA sequences and structures]] Main articles: Molecular biology, Cell biology, Genetics, Developmental biology Molecular biology is the study of biology at a molecular level. This field overlaps with other areas of biology, particularly with genetics and biochemistry. Molecular biology chiefly concerns itself with understanding the interactions between the various systems of a cell, including the interrelationship of DNA, RNA, and protein synthesis and learning how these interactions are regulated. Cell biology studies the physiological properties of cells, as well as their behaviors, interactions, and environment. This is done both on a microscopic and molecular level. Cell biology researches both single-celled organisms like bacteria and specialized cells in multicellular organisms like humans. Understanding cell composition and how they function is fundamental to all of the biological sciences. Appreciating the similarities and differences between cell types is particularly important in the fields of cell and molecular biology. These fundamental similarities and differences provide a unifying theme, allowing the principles learned from studying one cell type to be extrapolated and generalized to other cell types. Genetics is the science of genes, heredity, and the variation of organisms. In modern research, genetics provides important tools in the investigation of the function of a particular gene, or the analysis of genetic interactions. Within organisms, genetic information generally is carried in chromosomes, where it is represented in the chemical structure of particular DNA molecules. Genes encode the information necessary for synthesizing proteins, which in turn play a large role in influencing (though, in many instances, not completely determining) the final phenotype of the organism. Developmental biology studies the process by which organisms grow and develop. Originating in embryology, modern developmental biology studies the genetic control of cell growth, differentiation, and "morphogenesis," which is the process that gives rise to tissues, organs, and anatomy. Model organisms for developmental biology include the round worm Caenorhabditis elegans, the fruit fly Drosophila melanogaster, the zebrafish Brachydanio rerio, the mouse Mus musculus, and the weed Arabidopsis thaliana.

Physiology of organisms

Main articles: Physiology, Anatomy Physiology studies the mechanical, physical, and biochemical processes of living organisms by attempting to understand how all of the structures function as a whole. The theme of "structure to function" is central to biology. Physiological studies have traditionally been divided into plant physiology and animal physiology, but the principles of physiology are universal, no matter what particular organism is being studied. For example, what is learned about the physiology of yeast cells can also apply to human cells. The field of animal physiology extends the tools and methods of human physiology to non-human species. Plant physiology also borrows techniques from both fields. Anatomy is an important branch of physiology and considers how organ systems in animals, such as the nervous, immune, endocrine, respiratory, and circulatory systems, function and interact. The study of these systems is shared with medically oriented disciplines such as neurology and immunology.

Diversity and evolution of organisms

immunology of a population of organisms is sometimes depicted as if travelling on a fitness landscape. The arrows indicate the preferred flow of a population on the landscape, and the points A, B, and C are local optima. The red ball indicates a population that moves from a very low fitness value to the top of a peak]] Main articles: Evolutionary biology, Botany, Zoology Evolutionary biology is concerned with the origin and descent of species, as well as their change over time, and includes scientists from many taxonomically-oriented disciplines. For example, it generally involves scientists who have special training in particular organisms such as mammalogy, ornithology, or herpetology, but use those organisms as systems to answer general questions about evolution. Evolutionary biology also makes use of paleontologists, who use the fossil record to answer questions about the mode and tempo of evolution, as well as theoreticians in areas such as population genetics and evolutionary theory. In the 1990s, developmental biology re-entered evolutionary biology from its initial exclusion from the modern synthesis through the study of evolutionary developmental biology. Related fields which are often considered part of evolutionary biology are phylogenetics, systematics, and taxonomy. The two major traditional taxonomically-oriented disciplines are botany and zoology. Botany is the scientific study of plants. Botany covers a wide range of scientific disciplines that study the growth, reproduction, metabolism, development, diseases, and evolution of plant life. Zoology involves the study of animals, including the study of their physiology within the fields of anatomy and embryology. The common genetic and developmental mechanisms of animals and plants is studied in molecular biology, molecular genetics, and developmental biology. The ecology of animals is covered under behavioral ecology and other fields.

Classification of life

The dominant classification system is called Linnaean taxonomy, which includes ranks and binomial nomenclature. How organisms are named is governed by international agreements such as the International Code of Botanical Nomenclature (ICBN), the International Code of Zoological Nomenclature (ICZN), and the International Code of Nomenclature of Bacteria (ICNB). A fourth Draft BioCode was published in 1997 in an attempt to standardize naming in these three areas, but it has yet to be formally adopted. The International Code of Virus Classification and Nomenclature (ICVCN) remains outside the BioCode.

Interactions of organisms

International Code of Virus Classification and Nomenclature]] Main articles: Ecology, Ethology, Behavior, Biogeography Ecology studies the distribution and abundance of living organisms, and the interactions between organisms and their environment. The environment of an organism includes both its habitat, which can be described as the sum of local abiotic factors such as climate and geology, as well as the other the organisms that share its habitat. Ecological systems are studied at several different levels, from individuals and populations to ecosystems and the biosphere. As can be surmised, ecology is a science that draws on several disciplines. Ethology studies animal behavior (particularly of social animals such as primates and canids), and is sometimes considered a branch of zoology. Ethologists have been particularly concerned with the evolution of behavior and the understanding of behavior in terms of the theory of natural selection. In one sense, the first modern ethologist was Charles Darwin, whose book The expression of the emotions in animals and men influenced many ethologists. Biogeography studies the spatial distribution of organisms on the Earth, focusing on topics like plate tectonics, climate change, dispersal and migration, and cladistics.

History of the word "biology"

Formed by combining the Greek βίος (bios), meaning 'life', and λόγος (logos), meaning 'study of', the word "biology" in its modern sense seems to have been introduced independently by Gottfried Reinhold Treviranus (Biologie oder Philosophie der lebenden Natur, 1802) and by Jean-Baptiste Lamarck (Hydrogéologie, 1802). The word itself is sometimes said to have been coined in 1800 by Karl Friedrich Burdach, but it appears in the title of Volume 3 of Michael Christoph Hanov's Philosophiae naturalis sive physicae dogmaticae: Geologia, biologia, phytologia generalis et dendrologia, published in 1766.

History

Main articles: History of biology, History of medicine, History of genetics Major discoveries in biology include:
- Cell theory
- Germ theory of disease
- Genetics
- Evolution
- DNA

Related topics

Main articles: List of biology topics

External links


- [http://www.rom.on.ca/biodiversity/biocode/biocode1997.html BioCode]: A proposal for organism naming.
- [http://www.ncbi.nlm.nih.gov/entrez/query.fcgi?db=Books NCBI Open-Access Books]
- PhyloCode, [http://www.ohiou.edu/phylocode/index.html]
- [http://tolweb.org/tree/phylogeny.html The Tree of Life]: A multi-authored, distributed Internet project containing information about phylogeny and biodiversity.
- [http://www.bioone.org/perlserv/?request=index-html BioOne] Bioscience research journals.
- [http://www.bionews.in/biologynews.htm Biology News] Biology News, Articles and Research discoversies.

Further reading


- Lynn Margulis, Five Kingdoms: An Illustrated Guide to the Phyla of Life on Earth, 3rd ed., St. Martin's Press, 1997, paperback, ISBN 0805072527 (many other editions)
- Neil Campbell, Biology (7th edition), Benjamin-Cummings Publishing Company, 2004, hardcover, ISBN 080537146X
- Category:School subjects als:Biologie ko:생물학 ms:Biologi ja:生物学 simple:Biology th:ชีววิทยา

Wikipedia:Biology basic topics

See List of biology topics for the complete alphabetical list of Wikipedia articles addressing topics in biology. Below is a structured list of only the basic topics. Please see the most basic encyclopedia article topics for general instructions on constructing this list, and consult complete list of encyclopedia topics. Structured list :

Ecology & Evolution


- Life: origin of life -- Miller-Urey experiment
- Ecology:
  - Autecology: autotroph -- heterotroph -- acclimitization -- endotherm -- ectotherm -- hibernation -- homeostasis -- behavior -- circadian rhythm
  - Population ecology: population -- competition -- mating -- biological dispersal -- endemism -- niche -- growth curve -- carrying capacity
  - Community ecology: community -- keystone species -- mimicry -- symbiosis -- pollination -- mutualism -- commensalism -- parasitism -- predation -- invasive species -- environmental heterogeneity -- edge effect
  - Ecosystems: biodiversity -- biome -- habitat -- plankton -- thermocline -- carbon cycle -- water cycle -- nitrogen cycle -- food web -- trophic level -- saprobe -- decomposition
- Evolutionary biology (evolution)
  - Microevolution: species -- speciation -- adaptation -- selection -- natural selection -- directional selection -- sexual selection -- genetic drift -- sexual reproduction -- asexual reproduction -- colony -- allele frequency -- neutral theory of molecular evolution -- population genetics -- Hardy-Weinberg principle
  - Macroevolution: adaptive radiation -- convergent evolution -- extinction -- mass extinction -- fossil -- taphonomy -- geologic time -- plate tectonics -- continental drift -- vicariance -- Gondwana -- Pangaea -- endosymbiosis
  - Systematics: taxon -- taxonomy -- scientific classification -- phylogeny -- evolutionary tree -- cladistics -- synapomorphy -- homology -- molecular clock

Organismal biology


- Groups of living organisms
  - viruses : DNA viruses -- RNA viruses -- retroviruses
  - Single-cell organisms:
    - prokaryotes : microbe -- bacteria -- archaebacteria -- eubacteria
    - eukaryotes: fungi -- algae -- protozoa -- protista
  - Multicellular organisms:
    - plantae -- plants -- bryophytes -- pteridophytes -- seed plants
    - animalia -- animals -- metazoa -- insects -- molluscs -- vertebrates
    - fungi -- lichen -- mycorrhizae
- Developmental biology : gamete -- spermatid -- ovum -- zygote -- embryo -- cellular differentiation -- morphogenesis --homeobox
  - Animal development: stem cell -- blastula -- gastrula -- egg (biology) -- fetus -- placenta
  - Plant development: seed -- cotyledon -- meristem -- apical meristem -- vascular cambium -- cork cambium
- Morphology, Anatomy, & Physiology:
  - Techniques: electrophysiology -- electron microscopy
  - tissues -- organ -- senescence
  - Plant systems: root -- shoot -- stem -- leaf -- flower
    - vascular tissue -- Casparian strip -- turgor pressure -- xylem -- phloem -- transpiration -- wood
    - alternation of generations -- gametophyte -- antheridium -- archegonium -- sporophyte -- spore -- sporangium
    - tropism -- taxis
  - Animal systems: skin cell
    - skeleton -- bone -- cartilage -- joint -- muscle -- tendon -- actin -- myosin -- reflex
    - circulatory system -- respiration -- lung -- heart -- artery -- vein -- capillary -- blood -- blood cell
    - digestive system -- stomach -- intestine -- liver -- nutrition -- primary nutritional groups metabolism -- kidney -- excretion
    - nervous system: limbic system -- vestibular system -- neuron -- axon -- dendrite -- brain -- eye -- vision -- audition -- proprioception -- olfaction -- memory
    - nerve signaling: action potential -- signal transduction -- synapse -- receptor
    - endocrine system -- hormone
    - reproductive system -- testes -- ovary -- pregnancy
- Medicine:
  - Diseases: cancer -- diabetes -- obesity -- hereditary disease -- communicable disease
  - immune system: antibody -- host -- vaccine -- immune cell -- AIDS -- T-cell -- leucocyte
  - Drugs: antibiotic -- barbiturate -- SSRI
  - Techniques: MRI -- Computed axial tomography -- blood count

Cell & Molecular biology


- Cell biology: the cell
  - Techniques: culture -- microscope -- SEM -- TEM
  - cell components:
    - organelles: Golgi apparatus -- nucleus -- cytoplasm -- vacuole -- vesicle -- lysosome -- peroxisome -- mitochondrion -- plastid -- chloroplast
    - membranes: plasma membrane -- endomembrane system -- endoplasmic reticulum -- mitochondrial membrane -- chloroplast membrane
    - Other subcellular features: cell wall -- pseudopod -- cytoskeleton -- mitotic spindle -- flagellum -- cilium
  - cell processes:
    - diffusion -- osmosis -- isotonic -- active transport -- phagocytosis
    - energy pathways: fermentation -- glycolysis -- cellular respiration -- citric acid cycle -- photosynthesis -- Calvin cycle
    - protein biosynthesis -- ribosomes
    - cellular reproduction: cytokinesis -- centromere -- meiosis
    - nuclear reproduction: mitosis -- interphase -- prophase -- metaphase -- anaphase -- telophase
    - programmed cell death -- apoptosis -- cell senescence
- Biochemistry :
  - Biomolecules:
    - Small: amino acids -- Adenosine triphosphate (ATP) -- fats -- lipids -- oil -- phospholipid -- prion -- sugar -- vitamins -- neurotransmitter -- wax
    - Medium-sized:
      - pigments: chlorophyll -- carotenoid -- xanthophyll -- melanin
    - Biopolymers/macromolecules
      - proteins: primary structure -- secondary structure -- tertiary structure -- conformation -- native state -- protein folding -- enzyme -- receptor -- transmembrane receptor -- ion channel -- membrane transporter-- collagen -- silk
      - DNA -- RNA
      - polysaccharide: cellulose -- carbohydrate -- chitin -- glycogen -- starch
  - Biochemical mechanisms: proteolysis -- cooperativity
  - Techniques: electrophoresis -- His tag --affinity chromatography -- x-ray diffraction -- Proteomics -- mass spectrometry
- Molecular biology:
  - genetic structure: DNA -- DNA replication -- nucleosome -- genetic code -- codon -- transcription factor -- transcription -- translation -- RNA -- histone -- telomere
  - gene expression -- heterochromatin -- promoter -- enhancer -- operon
  - mutation -- point mutation -- crossover -- recombination --plasmid -- transposon
  - molecular genetics: DNA fingerprinting -- genetic fingerprint -- microsatellite -- gene knockout-- imprinting -- RNA interference
  - Genomics: computational biology -- bioinformatics
  - Molecular Techniques: gel electrophoresis -- transformation -- PCR -- PCR mutagenesis -- primer -- chromosome walking -- RFLP -- restriction enzyme -- sequencing -- shotgun sequencing -- cloning -- culture -- DNA microarray
- Genetics (classical genetics) :
  - heredity -- Mendelian inheritance -- gene -- locus -- trait -- allele -- polymorphism -- homozygote -- heterozygote -- hybrid -- hybridization -- dihybrid cross -- Punnett square
  - genotype-phenotype distinction -- genotype -- phenotype -- dominant gene -- recessive gene
  - genetic interactions -- segregation -- genetic mosaic -- maternal effect -- penetrance -- complementation -- suppression -- epistasis -- genetic linkage
  - chromosomal effects: chromosome -- haploid -- diploid -- polyploidy -- dosage effect -- inbreeding
  - Model organisms: Drosophila -- Arabidopsis -- Caenorhabitis elegans -- mouse -- Saccharomyces cerevisiae -- Escherichia coli -- Lambda phage -- Xenopus
  - Techniques: genetic screen -- paternity test -- linkage map -- genetic map

Biological disciplines


- Biology divisions : Acarology -- Anatomy -- Arachnology -- Biochemistry -- Bioinformatics -- Biomechanics -- Bionomics -- Biophysics -- Biotechnology -- Botany -- Cell biology -- Ecology -- Entomology -- Evolution -- Genetics -- Gerontology -- Herpetology -- Histology -- Ichnology -- Ichthyology -- Immunology -- Limnology -- Marine biology -- Microbiology -- Molecular biology -- Mycology -- Myrmecology -- Neurobiology -- Ornithology -- Paleontology -- Palynology -- Parasitology -- Photobiology -- Phycology -- Physiology -- Plant physiology -- Biological psychology-- Radiobiology -- Sociobiology -- Structural biology -- Taxonomy -- Virology -- Zoology
- Biologists: Charles Robert Darwin -- Alfred Russel Wallace -- Jean-Baptiste Lamarck -- Sir Charles Lyell -- Alfred Wegener -- Alexander Fleming -- Andrew Huxley -- Robert Koch -- Konrad Lorenz -- Ernst Haeckel -- Theodor Bilharz -- Marcello Malpighi -- Antoni van Leeuwenhoek -- Konrad Lorenz -- Thomas Malthus -- Edward Jenner -- Carolus Linnaeus -- Sewall Wright -- Louis Pasteur -- Gregor Mendel -- Barbara McClintock -- James D. Watson -- Francis Crick -- Kary Mullis -- Craig Venter -- Steven Jay Gould -- Lynn Margulis -- Carl Woese -- Jane Goodall Biology Biology Biology



Life

:For other uses, see Life and Living Life is a multi-faceted concept. Life may refer to the ongoing process of which living things are a part, the period between the conception (or a point at which the entity can be considered to be an individualized being) and death of an organism, the condition of an entity that has been born (or reached the point in its existence at which it can be established to be alive) and has yet to die, and that which makes a living thing alive.

Defining the concept of life

How can one tell when an entity is a lifeform? It would be relatively straightforward to offer a practical set of guidelines if one's only concern were life on Earth as we know it (see biosphere), but as soon as one considers questions about life's origins on Earth, or the possibility of extraterrestrial life, or the concept of artificial life, it becomes clear that the question is fundamentally difficult and comparable in many respects to the problem of defining intelligence. Also, loosely speaking, some theories are grounded in the basic assumption that "ideas have a life of their own".

A conventional definition

In biology, a lifeform has traditionally been considered to be a member of a population whose members can exhibit all the following phenomena at least once during their existence: #Growth, full development, maturity #Metabolism, consuming, transforming and storing energy/mass; growing by absorbing and reorganizing mass; excreting waste #Motion, either moving itself, or having internal motion #Reproduction, the ability to create entities that are similar to, yet separate from, itself or consisting solely of entities that exhibit the quality of reproduction. #Response to stimuli - the ability to measure properties of its surrounding environment, and act upon certain conditions. This property is also called homeostasis.

Exceptions to the conventional definition

These criteria are not without their uses, but their disparate nature makes them unsatisfactory from a number of perspectives; in fact, it is not difficult to find counterexamples and examples that require further elaboration. For example, according to the above definition, one could say:
- (most) mules and people who are infertile cannot reproduce and thus would not qualify as lifeforms. Also worker bees and other organisms living in colonies would not qualify; only the queen and the drones (or the whole colony) can be considered 'alive'.
- Fire and stars could be considered lifeforms.
- A virus does not grow and cannot reproduce outside of a host cell and thus would not qualify as a lifeform. Many individual organisms are incapable of reproduction and yet are still considered to be lifeforms; see mules and ants for examples. This is because the term "lifeform" applies on the level of entire species or of individual genes. (For example, see kin selection for information about one way by which non-reproducing individuals can still enhance the spread of their genes and the survival of their species.) It is important to keep in mind the difference between a "lifeform" and "a being that is alive." One example of sterility does not render the rest of the species a non-lifeform, any more than one dead animal renders the rest of the species dead. Note also that the two cases of fire and stars fitting the definition of life can be simply remedied by defining metabolism in a more biochemically exact way. Fundamentals of Biochemistry by Donald Voet and Judith Voet (ISBN 0471586501) defines metabolism as follows: "Metabolism is the overall process through which living systems acquire and utilize the free energy they need to carry out their various functions. They do so by coupling the exergonic reactions of nutrient oxidation to the endergonic processes required to maintain the living state, such as the performance of mechanical work, the active transport of molecules against concentration gradients, and the biosynthesis of complex molecules." This definition, in use by most biochemists, makes it clear that fire is not alive, because fire releases all the oxidative energy of its fuel as heat. (Note: Actually, the definition does not help much at all, for it is circular. What we are looking for, after all, is a definition of "living entity." We agreed that part of the definition is "capable of metabolism." We then tried to define "metabolism" in order to get clear on which entities are capable of it and which not. But the definition of "metabolism" just offered is in terms of living systems, and those are exactly what we are trying to define!) This could also be remedied by adding the requirement of locality, where there is an obvious structure that delineates the spatial extension of the living being, such as a cell membrane. A conceptual problem with saying that fire is life is that it collapses the distinction between "growth" and "reproduction." It is possible to think of a spreading flame as either growing or reproducing, but what would it mean to say that the same act is both growth and reproduction? Viruses reproduce, flames grow, some software programs mutate and evolve, future software programs will probably evince (even high-order) behavior, machines move, and some form of proto-life consisting of metabolizing cells without the ability to reproduce presumably existed. Still, some would not call these entities alive. Generally, all five characteristics are required for a population to be considered a lifeform.

Other definitions

Biologists who are content to focus on terrestrial organisms often note some additional signs of life, including these: # Living organisms contain molecular components such as: carbohydrates, lipids, nucleic acids, and proteins. # Living organisms require both energy and matter in order to continue living. # Living organisms are composed of at least one cell. # Living organisms maintain homeostasis for some period of time. # Species of living organisms will evolve. All life on Earth is based on the chemistry of carbon compounds. Some assert that this must be the case for all possible forms of life throughout the universe; others describe this position as 'carbon chauvinism'. The systemic definition is that living things are self-organizing and autopoietic (self-producing). These objects are not to be confused with dissipative structures (e.g. fire). Variations of this definition include:
- Francisco Varela and Humberto Maturana's definition of life (also widely used by Lynn Margulis) as an autopoietic (self-producing), water based, lipid-protein bound, carbon metabolic, nucleic acid replicated, protein readout system
- "a system of inferior negative feedbacks subordinated to a superior positive feedback" ([http://www.mol.uj.edu.pl/~benio/cyber_def_life.pdf J. theor Biol. 2001])
- Tom Kinch's definition of life as a highly organized auto-cannibalizing system naturally emerging from conditions common on planetary bodies, and consisting of a population of replicators capable of mutation, around each set of which a homeostatic metabolizing organism, which actively helps reproduce and/or protect the replicator(s), has evolved
- Stuart Kauffman's definition of life as an autonomous agent or a multi-agent system capable of reproducing itself or themselves, and of completing at least one thermodynamic work cycle
- Robert Pirsig's definition of life, found in his book Lila: An Inquiry into Morals, as that which maximizes its range of possible futures, in other words, that which makes decisions that result in the most future choices, or that which strives to keep its options open.
- A system converting entropy to negentropy, using flow of energy. Other definitions:
- That which seeks to continue its own existence (attributed to Clifford A. Schaffer).
- A self-replicating system that evolves through mutation.

Descent with modification: a "useful" characteristic

A useful characteristic upon which to base a definition of life is that of descent with modification: the ability of a life form to produce offspring that are like its parent or parents, but with the possibility of some variation due to chance. Descent with modification is sufficient by itself to allow evolution, assuming that the variations in the offspring allow for differential survival. The study of this form of heritability is called genetics. In all known life forms (assuming prions are not counted as such), the genetic material is primarily DNA or the related molecule, RNA. Another exception might be the software code of certain forms of viruses and programs created through genetic programming, but whether computer programs can be alive even by this definition is still a matter of some contention.

Origin of life

Main article: Origin of life There is no truly "standard" model of the origin of life, but most currently accepted scientific models build in one way or another on the following discoveries, which are listed roughly in order of postulated emergence: #Plausible pre-biotic conditions result in the creation of the basic small molecules of life. This was demonstrated in the Urey-Miller experiment. #Phospholipids spontaneously form lipid bilayers, the basic structure of a cell membrane. #Procedures for producing random RNA molecules can produce ribozymes, which are able to produce more of themselves under very specific conditions. There are many different hypotheses regarding the path that might have been taken from simple organic molecules to protocells and metabolism. Many models fall into the "genes-first" category or the "metabolism-first" category, but a recent trend is the emergence of hybrid models that do not fit into either of these categories.

The possibility of extraterrestrial life

Main articles: Extraterrestrial life, Astrobiology As of 2005, Earth is the only planet in the universe known by humans to support life. The question of whether life exists elsewhere in the universe remains open, but analyses such as the Drake equation have been used to estimate the probability of such life existing. There have been a number of claims of the discovery of life elsewhere in the universe, but none of these have yet survived scientific scrutiny. Today, the closest that scientists have come to finding extraterrestrial life is fossil evidence of possible bacterial life on Mars (via the ALH84001 meteorite). Searches for extraterrestrial life are currently focusing on planets and moons believed to possess liquid water, at present or in the past. Recent evidence from the NASA rovers Spirit and Opportunity supports the theory that Mars once had surface water. See Life on Mars for further discussion. Jupiter's moons are also considered good candidates for extraterrestrial life, especially Europa, which seems to possess oceans of liquid water. Other highly speculative and somewhat doubtful places for present or past life include the atmosphere of Venus, Titan cryovolcanoes, or even Enceladus.

See also


- Animal
- Artificial life
- Bacteria
- Biology
- Death
- Fungi
- Biological kingdom
- Biological life cycle
- Monera
- Odic force
- Origin of life (disambiguation)
- Plant
- Prehistoric life
- Protista

References


- Kauffman, Stuart. The Adjacent Possible: A Talk with Stuart Kauffman. Retrieved Nov. 30, 2003 from [http://www.edge.org/3rd_culture/kauffman03/kauffman_index.html]

External links


- [http://www.lifetheory.com Express your theory and meaning of life]
- [http://www.edge.org/3rd_culture/kauffman03/kauffman_index.html "The Adjacent Possible: A Talk with Stuart Kauffman"]
- [http://www.quotesandpoem.com/poems/SelectedPoetryTopic/Life Poems and Quotes about life and living]
- [http://www.angelfire.com/linux/vjtorley/ Animals and other living things: their interests, mental capacities and moral entitlements]
- [http://tolweb.org/tree?group=life Tree of Life Web Project - Life on Earth]
- [http://plato.stanford.edu/entries/life/ Stanford Encyclopedia of Philosophy entry]
- [http://web.archive.org/web/20041030074958/http://people.cornell.edu/pages/tg21/DHB.html The Deep Hot Biosphere Theory (Thomas Gold)] Category:Biology ja:生命 ko:생명 ms:Benda hidup simple:Life

Behavior

Behavior (or behaviour) refers to the actions or reactions of an object or organism, usually in relation to the environment. Behavior can be conscious or unconscious, overt or covert, and voluntary or involuntary. Behavior is controlled by the endocrine system, and the nervous system. The complexity of the behavior of an organism is related to the complexity of its nervous system. Generally, organisms with complex nervous systems have a greater capacity to learn new responses and thus adjust their behavior. of people (and other organisms or even mechanisms) falls within a range with some behaviors being common, some unusual, some acceptable, and some outside acceptable limits. The acceptablity of behavior is evaluated relative to social norms and regulated by various means of social control. For behavior of people see human behavior. In sociology, behavior is considered as having no meaning, being not directed at other people and thus is the most basic human action. Behavior should not be mistaken with social behavior, which is more advanced action, as social behavior is behavior specifically directed at other people. Animal behavior is studied in comparative psychology, ethology, behavioral ecology and sociobiology.

See also


- Ethology (animal behavior)
- Behaviorism
- Behavioral economics
- Behavior-based_robotics
- Chaining
- Deviant behavior
- Eccentricity (behaviour)
- Forms of activity and interpersonal relations
- Human behavior
- Instinct
- Normal (behavior)
- Reasoning
- Taboo
- Work behavior

External link


- [http://www.colorado.edu/epob/epob3730rlynch/01introduction.html Brain and behavior – (EPOB 3730) - University of Colorado]
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Species

In biology, a species is the basic unit of biodiversity. In scientific classification, a species is assigned a two-part name in Latin. The genus is listed first (and capitalized), followed by a specific epithet. For example, humans belong to the genus Homo, and are in the species Homo sapiens. The name of the species is the whole binomial not just the second term (the specific epithet). The binomial, and most other purely formal aspects of the biological codes of nomenclature, were formalized by Carolus Linnaeus in the 1700's and as a result are called the "Linnaean system". At that time, species were thought to represent independent acts of creation by God, and were therefore considered objectively real and immutable. Since the advent of the theory of evolution, the conception of species has undergone vast changes in biology, however no consensus on the definition of the word has yet been reached. The most commonly cited definition of "species" was first coined by Ernst Mayr. By this definition, called the biological species concept or isolation species concept, species are "groups of actually or potentially interbreeding natural populations which are reproductively isolated from other such groups". However, many other species concepts are also used (see other definitions of species below). The scientific name of a species is properly typeset in italics. When an unknown species is being referred to this may be done by using the abbreviation "sp." in the singular or "spp." in the plural in the place of the second part of the scientific name. Note that the word "specie" is not the singular of "species". It refers to coined money.

Definitions of species

The definition of a species given above as taken from Mayr, is somewhat idealistic. Since it assumes sexual reproduction, it leaves the term undefined for a large class of organisms that reproduce asexually. Biologists frequently do not know whether two morphologically similar groups of organisms are "potentially" capable of interbreeding. Further, there is considerable variation in the degree to which hybridization may succeed under natural and experimental conditions, or even in the degree to which some organisms use sexual reproduction between individuals to breed. Consequently, several lines of thought in the definition of species exist: ; Typological species : A group of organisms in which individuals are members of the species if they sufficiently conform to certain fixed properties. The clusters of variations or phenotypes within specimens (ie: longer and shorter tails) would differentiate the species. This method was used as a "classical" method of determining species, such as with Linnaeus early in evolutionary theory. However, we now know that different phenotypes do not always constitute different species (e.g.: a 4-winged Drosophila born to a 2-winged mother is not a different species). Species named in this manner are called morphospecies. ; Morphological species : A population or group of populations that differs morphologically from other populations. For example, we can distinguish between a chicken and a duck because they have different shaped bills and the duck has webbed feet. Species have been defined in this way since well before the beginning of recorded history. This species concept is much criticised because more recent genetic data reveals that genetically distinct populations may look very similar and, contrarily, large morphological differences sometimes exist between very closely-related populations. Nonetheless, most species known have been described solely from morphology. ; Biological / Isolation species : A set of actually or potentially interbreeding populations. This is generally the most useful formulation for scientists working with living examples of the higher taxa like mammals, fish, and birds, but meaningless for organisms that do not reproduce sexually. It does not distinguish between the theoretical possibility of interbreeding and the actual likelihood of gene flow between populations and is thus impractical in instances of allopatric (geographically isolated) populations. The results of breeding experiments done in artificial conditions may or may not reflect what would happen if the same organisms encountered each other in the wild, making it difficult to gauge whether or not the results of such experiments are meaningful in reference to natural populations. ; Mate-recognition species : A group of organisms that are known to recognise one another as potential mates. Like the isolation species concept above, it applies only to organisms that reproduce sexually. Unlike the isolation species concept, it focuses specifically on pre-mating reproductive isolation. ; Phylogenetic / Evolutionary / Darwinian species : A group of organisms that shares an ancestor; a lineage that maintains its integrity with respect to other lineages through both time and space. At some point in the progress of such a group, members may diverge from one another: when such a divergence becomes sufficiently clear, the two populations are regarded as separate species. ; Microspecies : Species that reproduce without meiosis or mitosis so that each generation is genetically identical to the previous generation. See also apomixis. In practice, these definitions often coincide, and the differences between them are more a matter of emphasis than of outright contradiction. Nevertheless, no species concept yet proposed is entirely objective, or can be applied in all cases without resorting to judgement. Given the complexity of life, some have argued that such an objective definition is in all likelihood impossible, and biologists should settle for the most practical definition. For most vertebrates, this is the biological species concept, and to a lesser extent (or for different purposes) the phylogenetic species concept. Many BSC subspecies are considered species under the PSC; the difference between the BSC and the PSC can be summed up insofar as that the BSC defines a species as a consequence of manifest evolutionary history, while the PSC defines a species as a consequence of manifest evolutionary potential. Thus, a PSC species is "made" as soon as an evolutionary lineage has started to separate, while a BSC species starts to exist only when the lineage separation is complete.

Importance in biological classification

The idea of species has a long history. It is one of the most important levels of classification, for several reasons:
- It often corresponds to what lay people treat as the different basic kinds of organism - dogs are one species, cats another.
- It is the standard binomial nomenclature (or trinomial nomenclature) by which scientists typically refer to organisms.
- It is the only taxonomic level which has empirical content, in the sense that asserting that two animals are of different species is saying something more than classificatory about them. After thousands of years of use, the concept remains central to biology and a host of related fields, and yet also remains at times ill-defined and controversial.

Implications of assignment of species status

The naming of a particular species should be regarded as a hypothesis about the evolutionary relationships and distinguishability of that group of organisms. As further information comes to hand, the hypothesis may be confirmed or refuted. Sometimes, especially in the past when communication was more difficult, taxonomists working in isolation have given two distinct names to individual organisms later identified as the same species. When two named species are discovered to be of the same species, the older species name is usually retained, and the newer species name dropped, a process called synonymization, or convivially, as lumping. Dividing a taxon into multiple, often new, taxons is called splitting. Taxonomists are often referred to as "lumpers" or "splitters" by their colleagues, depending on their personal approach to recognizing differences or commonalities between organisms (see lumpers and splitters). Traditionally, researchers relied on observations of anatomical differences, and on observations of whether different populations were able to interbreed successfully, to distinguish species; both anatomy and breeding behavior are still important to assigning species status. As a result of the revolutionary (and still ongoing) advance in microbiological research techniques, including DNA analysis, in the last few decades, a great deal of additional knowledge about the differences and similarities between species has become available. Many populations which were formerly regarded as separate species are now considered to be a single taxon, and many formerly grouped populations have been split. Any taxonomic level (species, genus, family, etc.) can be synonymized or split, and at higher taxonomic levels, these revisions have been still more profound. From a taxonomical point of view, groups within a species can be defined as being of a taxon hierarchically lower than a species. In zoology only the subspecies is used, while in botany the variety, subvariety, and form are used as well.

The isolation species concept in more detail

In general, for large, complex, organisms that reproduce sexually (such as mammals and birds), one of several variations on the isolation or biological species concept is employed. Often, the distinction between different species, even quite closely related ones, is simple. Horses (Equus caballus) and donkeys (Equus asinus) are easily told apart even without study or training, and yet are so closely related that they can interbreed after a fashion. Because the result, a mule or hinny, is not usually fertile, they are clearly separate species. But many cases are more difficult to decide. This is where the isolation species concept diverges from the evolutionary species concept. Both agree that a species is a lineage that maintains its integrity over time, that is diagnosably different to other lineages (else we could not recognise it), is reproductively isolated (else the lineage would merge into others, given the chance to do so), and has a working intra-species recognition system (without which it could not continue). In practice, both also agree that a species must have its own independent evolutionary history—otherwise the characteristics just mentioned would not apply. The species concepts differ in that the evolutionary species concept does not make predictions about the future of the population: it simply records that which is already known. In contrast, the isolation species concept refuses to assign the rank of species to populations that, in the best judgement of the researcher, would recombine with other populations if given the chance to do so.

The isolation question

There are, essentially, two questions to resolve. First, is the proposed species consistently and reliably distinguishable from other species? Secondly, is it likely to remain so in the future? To take the second question first, there are several broad geographic possibilities.
- The proposed species are sympatric—they occupy the same habitat. Observation of many species over the years has failed to establish even a single instance of two diagnostically different populations that exist in sympatry and have then merged to form one united population. Without reproductive isolation, population differences cannot develop, and given reproductive isolation, gene flow between the populations cannot merge the differences. This is not to say that cross breeding does not take place at all, simply that it has become negligible. Generally, the hybrid individuals are less capable of successful breeding than pure-bred individuals of either species.
- The proposed species are allopatric—they occupy different geographical areas. Obviously, it is not possible to observe reproductive isolation in allopatric groups directly. Often it is not possible to achieve certainty by experimental means either: even if the two proposed species interbreed in captivity, this does not demonstrate that they would freely interbreed in the wild, nor does it always provide much information about the evolutionary fitness of hybrid individuals. A certain amount can be inferred from other experimental methods: for example, do the members of population A respond appropriately to playback of the recorded mating calls of population B? Sometimes, experiments can provide firm answers. For example, there are seven pairs of apparently almost identical marine snapping shrimp (Altheus) populations on either side of the Isthmus of Panama, which did not exist until about 3 million years ago. Until then, it is assumed, they were members of the same seven species. But when males and females from opposite sides of the isthmus are placed together, they fight instead of mating. Even if the isthmus were to sink under the waves again, the populations would remain genetically isolated: therefore they are now different species. In many cases, however, neither observation nor experiment can produce certain answers, and the determination of species rank must be made on a 'best guess' basis from a general knowledge of other related organisms.
- The proposed species are parapatric—they have breeding ranges that abut but do not overlap. This is fairly rare, particularly in temperate regions. The dividing line is often a sudden change in habitat (an ecotone) like the edge of a forest or the snow line on a mountain, but can sometimes be remarkably trivial. The parapatry itself indicates that the two populations occupy such similar ecological roles that they cannot coexist in the same area. Because they do not crossbreed, it is safe to assume that there is a mechanism, often behavioral, that is preventing gene flow between the populations, and that therefore they should be classified as separate species.
- There is a hybrid zone where the two populations mix. Typically, the hybrid zone will include representatives of one or both of the 'pure' populations, plus first-generation and back-crossing hybrids. The strength of the barrier to genetic transmission between the two pure groups can be assessed by the width of the hybrid zone relative to the typical dispersal distance of the organisms in question. The dispersal distance of oaks, for example, is the distance that a bird or squirrel can be expected to carry an acorn; the dispersal distance of Numbats is about 15 kilometres, as this is as far as young Numbats will normally travel in search of vacant territory to occupy after leaving the nest. The narrower the hybrid zone relative to the dispersal distance, the less gene flow there is between the population groups, and the more likely it is that they will continue on separate evolutionary paths. Nevertheless, it can be very difficult to predict the future course of a hybrid zone; the decision to define the two hybridizing populations as either the same species or as separate species is difficult and potentially controversial.
- The variation in the population is clinal; at either extreme of the population's geographic distribution, typical individuals are clearly different, but the transition between them is seamless and gradual. For example, the Koalas of northern Australia are clearly smaller and lighter in colour than those of the south, but there is no particular dividing line: the further south an individual Koala is found, the larger and darker it is likely to be; Koalas in intermediate regions are intermediate in weight and colour. In contrast, over the same geographic range, black-backed (northern) and white-backed (southern) Australian Magpies do not blend from one type to another: northern populations have black backs, southern populations white backs, and there is an extensive hybrid zone where both 'pure' types are common, as are crossbreeds. The variation in Koalas is clinal (a smooth transition from north to south, with populations in any given small area having a uniform appearance), but the variation in magpies is not clinal. In both cases, there is some uncertainty regarding correct classification, but the consensus view is that species rank is not justified in either. The gene flow between northern and southern magpie populations is judged to be sufficiently restricted to justify terming them subspecies (not full species); but the seamless way that local Koala populations blend one into another shows that there is substantial gene flow between north and south. As a result, experts tend to reject even subspecies rank in this case.

The difference question

Obviously, when defining a species, the geographic circumstances become meaningful only if the populations groups in question are clearly different: if they are not consistently and reliably distinguishable from one another, then we have no grounds for believing that they might be different species. The key question in this context, is "how different is different?" and the answer is usually "it all depends". In theory, it would be possible to recognise even the tiniest of differences as sufficient to delineate a separate species, provided only that the difference is clear and consistent (and that other criteria are met). There is no universal rule to state the smallest allowable difference between two species, but in general, very trivial differences are ignored on the twin grounds of simple practicality, and genetic similarity: if two population groups are so close that the distinction between them rests on an obscure and microscopic difference in morphology, or a single base substitution in a DNA sequence, then a demonstration of restricted gene flow between the populations will probably be difficult in any case. More typically, one or other of the following requirements must be met:
- It is possible to reliably measure a quantitative difference between the two groups that does not overlap. A population has, for example, thicker fur, rougher bark, longer ears, or larger seeds than another population, and although this characteristic may vary within each population, the two do not grade into one another, and given a reasonably large sample size, there is a definite discontinuity between them. Note that this applies to populations, not individual organisms, and that a small number of exceptional individuals within a population may 'break the rule' without invalidating it. The less a quantitative difference varies within a population and the more it varies between populations, the better the case for making a distinction. Nevertheless, borderline situations can only be resolved by making a 'best-guess' judgement.
- It is possible to distinguish a qualitative difference between the populations; a feature that does not vary continuously but is either entirely present or entirely absent. This might be a distinctively shaped seed pod, an extra primary feather, a particular courting behaviour, or a clearly different DNA sequence. Sometimes it is not possible to isolate a single difference between species, and several factors must be taken in combination. This is often the case with plants in particular. In eucalypts, for example, Corymbia ficifolia cannot be reliably distinguished from its close relative Corymbia calophylla by any single measure (and sometimes individual trees cannot be definitely assigned to either species), but populations of Corymbia can be clearly told apart by comparing the colour of flowers, bark, and buds, number of flowers for a given size of tree, and the shape of the leaves and fruit. When using a combination of characteristics to distinguish between populations, it is necessary to use a reasonably small number of factors (if more than a handful are needed, the genetic difference between the populations is likely to be insignificant and is unlikely to endure into the future), and to choose factors that are functionally independent (height and weight, for example, should usually be considered as one factor, not two).

Historical development of the species concept

In the earliest works of science, a species was simply an individual organism that represented a group of similar or nearly identical organisms. No other relationships beyond that group were implied. Aristotle used the words genus and species to mean generic and specific categories. Aristotle and other pre-Darwinian scientists took the species to be distinct and unchanging, with an "essence", like the chemical elements. When early observers began to develop systems of organization for living things, they began to place formerly isolated species into a context. To the modern mind, many of the schemes delineated are whimsical at best, such as those that determined consanguinity based on color (all plants with yellow flowers) or behavior (snakes, scorpions and certain biting ants). In the 18th century Carolus Linnaeus classified organisms according to differences in the form of reproductive apparatus. Although his system of classification sorts organisms according to degrees of similarity, it made no claims about the relationship between similar species. At the time, it was still widely believed that there is no organic connection between species, no matter how similar they appear; every species was individually created by God, a view today called creationism. This approach also suggested a type of idealism: the notion that each species exists as an "ideal form". Although there are always differences (although sometimes minute) between individual organisms, Linnaeus considered such variation problematic. He strove to identify individual organisms that were exemplary of the species, and considered other non-exemplary organisms to be deviant and imperfect. By the 19th century most naturalists understood that species could change form over time, and that the history of the planet provided enough time for major changes. As such, the new emphasis was on determining how a species could change over time. Jean-Baptiste Lamarck suggested that an organism could pass on an acquired trait to its offspring, i.e., the giraffe's long neck was attributed to generations of giraffes stretching to reach the leaves of higher treetops (this well-known and simplistic example, however, does not do justice to the breadth and subtlety of Lamarck's ideas). Lamarck's most important insight may have been that species can be extraordinarily fluid; his 1809 Zoological Philosophy contained one of the first logical refutations of creationism. With the acceptance of the work of Charles Darwin in the 1860s, Lamarck's view of evolution was quickly eclipsed. It was not until the late 20th century that his work began to be reexamined, and took its place as a fundamental stepping stone to the modern theory of adaptive mutation. Lamarck's long-discarded ideas of the goal-oriented evolution of species, also known the teleological process, have also received renewed attention, particularly by proponents of artificial selection. Charles Darwin and Alfred Wallace provided what scientists now consider the most powerful and compelling theory of evolution. Basically, Darwin argued that it is populations that evolve, not individuals. His argument relies on a radical shift in perspective from Linnaeus: rather than defining species in ideal terms (and searching for an ideal representative and rejecting deviations), Darwin considered variation among individuals to be natural. He further argued that variation, far from being problematic, actually provides the explanation for the existence of distinct species. Darwin's work drew on Thomas Malthus' insight that the rate of growth of a biological population will always outpace the rate of growth of the resources in the environment, such as the food supply. As a result, Darwin argued, not all the members of a population will be able to survive and reproduce. Those that did will, on average, be the ones possessing variations—however slight—that make them slightly better adapted to the environment. If these variable traits are heritable, then the offspring of the survivors will also possess them. Thus, over many generations, adaptive variations will accumulate in the population, while counter-adaptive will be eliminated. It should be emphasized that whether a variation is adaptive or non-adaptive depends on the environment: different environments favor different traits. Since the environment effectively selects which organisms live to reproduce, it is the environment (the "fight for existence") that selects the traits to be passed on. This is the theory of evolution by natural selection. In this model, the length of a giraffe's neck would be explained by positing that proto-giraffes with longer necks would have had a significant reproductive advantage to those with shorter necks. Over many generations, the entire population would be a species of long-necked animals. In 1859, when Darwin published his theory of natural selection, the mechanism behind the inheritance of individual traits was unknown. Although Darwin made some speculations on how traits are inherited (pangenesis), his theory relies only on the fact that inheritable traits exist, and are variable (which makes his accomplishment even more remarkable.) Although Gregor Mendel's paper on genetics was published in 1866, its significance was not recognized. It was not until 1900 that his work was rediscovered by Hugo de Vries, Carl Correns and Erich von Tschermak, who realised that the "inheritable traits" in Darwin's theory are genes. The theory of the evolution of species through n